2Studies of the Actinoceratida
Part 1: The Ordiovician deveopment
of the Actinoceratida, with notes on Actinoceratida morphology and
Part 2: Macroloxeras, a Devoinian homeomorph
of the Actinoceratida
By R. H. Flower, 1957, 101 pp., 5 figs., 13 plates, 1 index.
Descriptions of new Ordovician species of Actinoceratida are combined with a review of the evolution and faunal development of the order in the Ordovician. New morphological information includes spatial relationships of cameral and siphonal deposits in complete shells of several species, and a revised concept of the canal system. Occurrences of antinoceroids in the Ordovician of North America, eastern Asia, and northern Europe are summarized. A revised concept of evolution of the group is presented. Reason is given for regarding Bathmoceras as the ancestor of the order; it gave rise to Polydesmia having thick primitive rings and dendritic radial canals. Reticulate canals characterize Whiterock actinoceroids, of which the genera Wutinoceras, Cyrtonybyoceras, and Adamsoceras are characteristic. Ormoceras, developed from Adamsoceras, continues beyond the close of the Ordovician, giving rise to Deiroceras, of which Troedssonoceras is a synonym. Actinoceras yields itself to division into species groups of faunal and stratigraphic value. Leurorthoceras is a synonym of this genus, and Saffordoceras and Troostoceras are doubtfully distinct. Kochoceras is derived from Actinoceras. Internal differences indicate that Gonioceras and Lambeoceras are homeorphic but not closely related.
Stratigraphic notes include a general discussion of the Ordovician, with special reference to the recent revisions, recognition of a pre-Chazyan interval, the Chazy-Black River hiatus, and the correlation of the Red River beds with the Cobourg and Eden. A revision of the Montoya Group is presented. The importance of austral-boreal oscillations is emphasized, the actinoceroids being predominantly boreal in faunal affinities, although in Cobourg time Deiroceras and Orthonybyoceras invaded the austral realm. The systematic portion includes discussion of 15 genera, one of which, Adamsoceras, is new. Thirty-one species are described and illustrated, all but two of which are new; these are listed in the table of contents.
The new genus Macroloxoceras is described from two species; the genotype M. magnum, from the Dyer Dolomite of Colorado, and M. minor, from the Percha Shale of New Mexico, both very late Devonian age. Together with Bergoceras, Paraloxoceras, and Pseudactinoceras, this genus comprises the new subfamily Macroloxoceratinae of the Pseudorthoceratidae. Although in the entire subfamily there is a strong resemblance to Rayonnoceras, reexaminations of the evidence suggests this relationship is homeomorphic, though a remarkable example of contemporaneous convergence. Pseudocyrtoceras, sp. Schindewolf is assigned to the Pseudorthoceratinae. In spite of its resemblance to the Macroloxoceratinae, Rayonnoceras retains actinoceroid features favoring its derivation from Ormoceras.
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